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Encyclopedia > Spiders
For other uses, see Spider (disambiguation).
Spiders

Long-jawed orb weaver spider
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Arachnida
Order: Araneae
Families

Suborder Mesothelae
    Liphistiidae (primitive burrowing spiders)
Suborder Mygalomorphae
    Atypidae (atypical tarantula)
    Antrodiaetidae (folding trapdoor spider)
    Mecicobothriidae (dwarf tarantulas)
    Hexathelidae (venomous funnel-web tarantula)
    Dipluridae (funnel-web tarantula)
    Cyrtaucheniidae (wafer trapdoor spider)
    Ctenizidae (trapdoor spider)
    Theraphosidae (tarantula)
Suborder Araneomorphae
    Hypochilidae (lampshade spider)
    Filistatidae (crevice weaver)
    Sicariidae (recluse spider)
    Scytodidae (spitting spider)
    Leptonetidae (leptonetid spider)
    Pholcidae (daddy long-legs spider)
    Plectreuridae (plectreurid spider)
    Diguetidae (coneweb spider)
    Caponiidae (two-eyed spider)
    Segestriidae (tube-dwelling spider)
    Dysderidae (woodlouse hunter spider)
    Oonopidae (oonopid spider)
    Palpimanidae (palp-footed spider)
    Mimetidae (pirate spider)
    Eresidae (velvet spider)
    Oecobiidae (including Urocteidae)
    (wall and six-exit tent spider)
    Hersiliiidae (tree trunk spider)
    Deinopidae (ogre-faced spider)
    Uloboridae (cribellate orb and
        sector weaver spider)
    Nesticidae (scaffold web spider)
    Theridiidae (tangle web spider)
    Linyphiidae (bowl and doily and
        dwarf spiders)
    Tetragnathidae (long jawed spider)
    Araneidae (orb-weaver spider)
    Lycosidae (wolf spider)
    Pisauridae (nursery web spider)
    Oxyopidae (lynx spider)
    Zorocratidae (zorocratid spider)
    Zoropsidae (zoropsid spider)
    Ctenidae (wandering spider)
    Agelenidae (araneomorph funnel-web spider)
    Cybaeidae (water spider)
    Desidae (intertidal spider)
    Hahniidae (dwarf sheet spider)
    Dictynidae (dictynid spider)
    Amaurobiidae (tangled nest spider)
    Titanoecidae (titanoecid spider)
    Tengellidae (tengellid spider)
    Miturgidae (long-legged sac spider)
    Anyphaenidae (anyphaenid sac spider)
    Liocranidae (liocranid sac spider)
    Clubionidae (sac spider)
    Corinnidae (corinnid sac spider)
    Zodariidae (zodariid ground spider)
    Gnaphosidae (ground spiders)
    Selenopidae (wall crab spider)
    Sparassidae (huntsman, etc.)
    Philodromidae (philodromid crab spider)
    Thomisidae (crab spider)
    Salticidae (jumping spider)
Source: Platnick 2003

Spiders are invertebrate animals that produce silk, have eight legs and no wings. More precisely, a spider is any member of the arachnid order Araneae, an order divided into three sub-orders in newer systems: the Mygalomorphae (the primitive spiders), the Araneomorphae (the modern spiders) and the Mesothelae, which contains the Family Liphistiidae, rarely seen burrowing spiders from Asia. The study of spiders is known as arachnology, although it is often grouped under the more general area of entomology.


Many spiders hunt by building webs to trap insects. These webs are made of spider silk, a thin, strong protein strand extruded by the spider from spinnerets on the end of the abdomen. All spiders produce silk, although not all use it to spin elaborate traps. Silk can be used to aid in climbing, forming smooth walls for burrows, cocooning prey, and for many other applications.

Contents

Anatomy

Spiders, unlike insects, have their bodies divided in only two segments: prosoma or cephalothorax (a fused head and thorax) and opisthosoma or abdomen. New World tarantulas have a patch of urticating hairs on their abdomens, while these are lacking in Old World species. Spiders have eight legs compared to the insects' six, and their eyes (usually eight) are single lenses rather than compound eyes like those of most insects.

Enlarge
A long-jawed spider illustrating jaws, pedipalps, and eye pattern

Eyes can be arranged differently in different species. Sometimes one pair is better developed than the rest. Some species have a pair less or are even without eyes. While several families of hunting spiders have good to excellent vision (Lycosidae and Salticidae), the members of most families of hunting spiders, most of the web weavers, and the spiders that lurk on flowers and other fixed locations waiting for some insect to come their way have very poor eyesight. Instead, these spiders hunt using their extreme sensitivity to vibrations.


Unlike insects, spiders have no antennae (feelers). Instead, they have pedipalps, sometimes just called palps, which are two additional appendages next to their mouth parts that, besides having other functions, they use to help themselves manipulate the food that they eat.


Spider haemolymph (the equivalent of blood) does not circulate through vessels, but just fills the body of the spider. This is called an open circulatory system. The spider's respiratory anatomy is based on a tracheal system, with each opening to the trachea as an extension of an outer pore reaching from the spider abdomen, protected by spiracles which are pores in the spider exoskeleton, composed of chitin.


Book lungs alone, respiratory organs with openings on the ventral surface of the abdomen, enrich the blood of primitive mygalomorph spiders with oxygen, and araneomorph spiders use spiracles as well. The latter are much more efficient and allow more advanced characteristics to be displayed, such as cursorial hunting (hunting involving rapid pursuit).


Life cycle

The spider life cycle progresses through three stages: the embryonic, the larval, and the nympho-imaginal (Foelix, 1996).


Between the time an egg is fertilized and the spider begins to take the shape of a spider is referred to as the embryonic stage (Foelix, 1996). As the spider begins to look more like a spider it enters the larval stage (Foelix, 1996). It enters the larval stage as a prelarva and, through subsequent molts, it reaches its larval form, a spider-looking, non self-sufficient animal feeding off its yolk supply (Foelix, 1996). After a few more molts, body structures become differentiated; all organ systems are complete and the animal begins to hunt on its own; it has reached the nympho-imaginal stage (Foelix, 1996). This stage is differentiated by two sub-stages: the nymph, or juvenile stage and the imago, or adult stage (Foelix, 1996). A spider does not transition from the nymph to the imago until it has become sexually mature (Foelix, 1996). Once a spider has reached the imago stage, it will remain here until its death.


Reproduction

Spiders reproduce by eggs laid in silk bundles called egg sacs. Spiders often use elaborate mating rituals (especially in the visually advanced jumping spiders) to allow the male to approach close enough to inseminate the female without triggering a predatory response. Assuming that the approach signals are exchanged correctly, the male spider must make a timely departure after mating to escape before the female's normal predatory instincts come back into operation.


Unusually, sperm transmission is an indirect process. When a male is ready to mate, he will spin a web pad onto which the contents of the abdominal reproductive organs are discharged. He then dips his palps (also known as 'palpi'), the small, leg-like appendages on the front of his cephalothorax, into the sperm, absorbing it. Mature male spiders characteristically have swollen bulbs on the end of their palps for this purpose, and this is a useful way to identify the sex of a spider in the field. With his palps thus 'charged' he then goes off in search of a female. The act of copulation occurs when the male inserts one or both palps into the fenale's genital opening, known as the epigyne. He transfers his sperm into the female by contracting his palps.


Very unusual behaviour is seen in spiders of the genus Tidarren, as the male amputates one of his palps before maturation and enters his adult life with one palp only. The palpi constitute 20% of its body mass, and since this weight greatly impedes its movement, the spider detaches one of the two to gain mobility. In the Yemani species Tidarren argo, the remaining palp is then torn off by the female. The separated palp remains attached to the female's epigynum for about four hours and apparently continues to function independently. In the meantime the female feeds on the palpless male. (Journal of Zoology (2001), 254:449-459 Cambridge University Press)


Do female spiders eat their mates?

It is often said that the male (usually significantly smaller than the female, down to 1% of her size for Tidarren sisyphoides) is likely to be killed by the female after the coupling, or sometimes before intercourse has occurred. This supposed propensity is what gave the black widow spider (Latrodectus mactans) its name, but it is the Australian red-backed spider (Latrodectus hasselti) that seems to require the sacrifice of the male.


The male Latrodectus hasselti actually dies after it inserts its second palpus in the female genital opening even if the female does not eat it. The three species of North American black widows do not seem to require the death of the male (although it may occur) and males can sometimes live in the web of a female for a while without being harmed.


However, despite these examples, and many other similar reports, the arresting story of the 'sacrificial male' has become perhaps somewhat greater than the truth. Mating of spiders is not invariably followed by cannibalism. Rainer F. Foelix, (1982), says "The supposed aggressiveness of the female spider towards the male is largely a myth... only in some exceptional cases does the male fall victim to the female.". Michael Roberts (1995) says "It is rare for a fit male to be eaten by the female..."


Possibly the general difficulty of observing spiders mating, and the wide variation in their mating habits, has allowed this idea to become so well-established. Undoubtedly there are occasions (as in Tidarren argo) when the female does eat the successful male suitor, and there has been much speculation on why this sacrifice of male mates might occur. One theory is that once the male has mated, if he is unlikely to mate again then further extension of his life serves no evolutionary purpose, while the sacrifice of the male may help increase egg production through increased nutrition for the female. Having more offspring would give the male an advantage of having his genes passed on over other males that were not eaten. This would be consistent with the hypothesis (Roberts, 1995) that old or unfit males get eaten, whilst younger and fitter ones may survive to mate again.


Ecology

Enlarge
Spider in web under rock

Spiders have a great range of variation and lifestyle, although all are predatory.


While spiders are generalist predators, in actuality their different methods of prey capture often limits the type of prey taken. Thus web-building spiders rarely capture caterpillars and crab spiders that ambush prey in flowers capture more bees, butterflies and some flies than other insects. Groups of families that tend to take certain types of prey because of their prey capture methods are often called guilds. A few spiders are more specialized in their prey capture. Dysdera captures and eats sowbugs, pillbugs and beetles, while pirate spiders eat only other spiders. Bolas spiders in the family Araneidae use sex pheromone analogs to capture only the males of certain moth species! Despite their generally broad prey ranges spiders are one of the most important links in the regulation of the populations of insects. Every day on a meadow they devour over 10 g/mē of insects and other arthropods.


Predatory techniques

Enlarge
A golden silk spider (Nephila clavipes), member of the family Tetragnathidae

There are many families of spiders, and the ways that they catch prey are diverse. But whether they catch insects, fish, small mammals, small birds, or some other small form of life, as soon as a spider makes contact with its prey it will attempt to bite.


Spiders bite their prey, and occasionally animals that cause them pain or threaten them, to do two things. First, they inflict mechanical damage, which, in the case of a spider that is as large or larger than its prey, can be severe. Second, they can choose to inject venom through their hollow fangs. Many genera, such as the widow spiders, inject neurotoxins that can spread through the prey's entire body and interfere with vital body functions. Other genera inject venom that operates to produce tissue damage at the site of the bite. Genera such as that of the brown recluse spider produce a necrotoxin. The necrotoxin is injected into prey where it causes the degradation of cell membranes. In the larger victims that do not die from these attacks, painful lesions over a fairly wide area of the body can remain active for fairly long periods of time.


Digestion is carried out internally and externally. The spiders secrete digestive fluids into their prey from a series of ducts perforating their jaws, These digestive fluids dissolve the prey's internal tissues.Then, the spider feeds by sucking the partially digested fluids out. Spiders consume only liquid foods. Many spiders will store prey temporarily while this process of external digestion is going on. Web weaving spiders that have made a shroud of silk to quiet their envenomed prey's death struggles will generally leave them in these shrouds and then consume them at their leisure.


Spider webs and prey capture

Main article: Spider web


Some spiders spin funnel-shaped webs, others make irregular webs, and still others make the spiral "orb" webs which are most commonly associated with the order.


The spider, after spinning its web, will then wait (often, but not always, at the center of the web) for a prey animal to become trapped. They sense the impact and struggle of a prey animal by vibrations transmitted along the web lines.


Other species of spiders do not use webs for capturing prey directly, instead pouncing from concealment (e.g. Trapdoor spiders) or running them down in open chase (e.g. Wolf spiders). Spiders do not usually adhere to their own webs. However, they are not immune to their own glue. Some of the strands of the web are sticky, and others are not. The spiders have to be careful to only climb on the non-sticky strands.


Types of spiders

Over 37,000 species of spiders have been identified but, because of their great ability for hiding, it is believed there are nearer 200,000 species. Almost all species are venomous, but only 30 species are known to be deadly dangerous.


Tangleweb spiders

Orb web spiders (Araneidae)

Other forms of webs

Spiders that live on flowers

Hunting spiders

Others

  • Kimura-gumo (Liphistiomorphae) (not dangerous)
  • Spruce-fir moss spider (Microhexura montivaga)
  • Tooth cave spider (Neoleptoneta myopica)

Spider bites

Most spiders are unlikely to bite humans because they do not identify humans as prey. Spiders, even small ones, may however bite humans when pinched. For instance, a common jumping spider (Family: Salticidae), around 3/8 inch (1 cm.) long, when pinched between the folds of a human's palm may inflict a bite that is about as painful as a bee sting. Many cold and temperate parts of the world, including the United Kingdom, are free from any spiders which are capable of inflicting any painful bite in normal circumstances: but in warmer climes there are a number of spiders which are capable of biting a human, and, of these, a few can also be classed as dangerous.


Dangerous spiders in the United States include widow spiders, brown recluse spiders, hobo spiders, and yellow sac spiders.


None of these spiders will intentionally "come after you," but they should be removed from one's house to avoid accidental injury. Many authorities warn against spraying poisons indiscriminately to kill all spiders, because doing so may actually remove one of the biological controls against incursions of the more dangerous species by ridding them of their competition.


If dangerous spiders are present in your area, then be mindful when you move cardboard boxes and other such objects that may have become the shelter of a poisonous spider. There is no need to be fearful, just do not grab a spider.


Black widows

Normally, black widow spider bites are fatal only to children, due to the fact that children have much smaller body weights than adults and so the poison is more concentrated in their bodies when a bite does occur.


There are several widow spiders, i.e., spiders of the genus Latrodectus, and they are generally regarded as all being about equally venomous. Widow spiders are practically blind, and move with difficulty when they are not on their web. Unlike the other problematical spiders, the females of these most toxic of U.S. spiders stay on their web, and the males (which wander around seeking mates) are too small to deliver a dangerous amount of venom.


The website of the Carlsen Ranch (http://www.nitehawk.com/CarlsenRanch/newsl/animal/spiders/htm) asserts that almost half the black widow bites reported in the medical literature from 1901-1941 "were inflicted on the male genitalia by spiders on the underside of outdoor toilet seats." From the standpoint of a spider that likes to make its web under overturned wooden boxes resting on the ground, and in other such sheltered places near the ground, and which eats flies, an outdoor toilet is an almost ideal location. If a black widow happens to be resting near the front edge of the hole when a male human visitor sits down, she may find herself pinned against the wood by his penis, and bite it.


Brown recluse spiders

Enlarge
Brown Recluse (Photo courtesy of the University of Nebraska)

Brown recluse spiders frequently wander about and so are more easily trapped against one's skin by clothing, bed sheets, etc. The so-called "aggressive house spider" or hobo spider (Tegenaria agrestis) is also a wanderer that may come into contact with people and bite. The yellow sac spiders take shelter in silk tubes during the daytime and come out to hunt at night. People may unintentionally make contact with them in the dark and so be bitten.


Brown recluse spider bites can produce very severe local symptoms, death of tissue around the wound, and, sometimes, severe systemic symptoms. The bites of hobo spiders can cause both pain and necrosis (tissue death), and yellow sac spider bites can be extremely painful.


Huntsman spiders

The huntsman spiders have a worldwide reputation for scaring people. They are large, defend their nests, and may move toward people and make threat displays. They frequently enter houses and hunt over the walls and ceilings where they may run rapidly for long distances without pausing. When they actually do bite people, the bites are very unpleasant, but these spiders are not regarded as dangerous.


There is one spider in California and Japan, probably a huntsman (tentatively identified as a member of the Sparassidae family, Heteropoda venatoria), that might run over and bite your finger if you touch the wall that it is clambering over. That behavior may well occur because its eyesight is good enough to see movement and general shape, but not sufficient to avoid mistaking something else for its natural prey.


Redback jumping spiders

Some people have reported being bitten by redback jumping spiders (Phidippus johnsoni). Most reports seem to be from California. These spiders have bright red abdomens (the females have a black stripe), and should be clearly visible even if they are not particularly large. It is unclear how the bites occur. Accidental contact seems rather unlikely since jumping spiders have excellent vision and could easily avoid being brushed by a human hand. It is also unlikely that they would mistake a human finger for their natural prey. One source suggests that since they are quite attractive children may try to pick them up and in that way elicit a defensive bite. Fortunately, the worst consequences reported have been three to four days of discomfort, with no permanent damage. Since they do not frequent human habitations it should ordinarily be easy to avoid unpleasant contact with them.


Brazilian wandering spiders and Sydney funnel-web spiders

The Brazilian wandering spider (a ctenid spider) and the Sydney funnel-web spider (a mygalomorph not related to the araneomorph funnel-web spiders) frequently bite people and are regarded as among the most dangerous in the world. The Sydney funnel-web spider is restricted to a relatively small area around Sydney, Australia. There are less dangerous species related to this spider in other parts of Australia, including Tasmania. The males in this case have somewhat more potent venom than females and they are also very aggressive. The Brazilian wandering spider reportedly may hitch a ride in clusters of bananas. As a result, any large spider appearing in a bunch of bananas should be treated with due care. Oddly many of the bites of this species are dry bites and no venom is released. Unfortunately if the venom is released the result can be very grave for the victim. The spiders are as large as some small tarantulas and have fairly long fangs.


Further reading

  • The World of Spiders, by W. S. Bristowe Collins (New Naturalist), London 1958
  • The Life of the Spider, by John Crompton. Mentor, 1950.
  • How to Know the Spiders, by B. J. Kaston. Dubuque, 1953.
  • Biology of Spiders, by Rainer F. Foelix, 1982 and second edition, 1996
  • The Book of the Spider, by Paul Hillyard, Random House, New York 1994
  • Spiders, by Barbara York Main, Collins (The Australian Naturalist Library), Sydney 1976
  • Spiders of Britain and Northern Europe, by Michael J. Roberts, Collins, London 1995

External links

  • Arachnology Home Pages: Araneae (http://www.arachnology.org/Arachnology/Pages/Araneae.html)
  • Spider info by Ed Nieuwenhuys (http://www.xs4all.nl/~ednieuw/Spiders/InfoNed/The_spider.html)
  • List of spiders found in Great Britain (http://www.britishspiders.org.uk/html/info.html)
  • Platnick, N.I. 2004. World Spider Catalog (http://research.amnh.org/entomology/spiders/catalog81-87/index.html)
  • Webs made by spiders fed on drug-dosed flies (http://www.cannabis.net/weblife.html)
  • Pictures of Spiders (http://www.surviveoutdoors.com/reference/spiders/index.asp)
  • Spider Bites (http://www.surviveoutdoors.com/emergency/spiderbites.asp)
  • University of California evaluations of spider bite severity (http://spiders.ucr.edu/dermatol.html)
  • Camel Spiders (http://www.camel-spiders.net)

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